Noun quill has 4 senses
  1. quill, quill pen - pen made from a bird's feather
    --1 is a kind of pen
  2. quill - a stiff hollow protective spine on a porcupine or hedgehog
    --2 is a kind of
    spine
    --2 is a part of
     porcupine, hedgehog; hedgehog, Erinaceus europaeus, Erinaceus europeaeus
  3. flight feather, pinion, quill, quill feather - any of the larger wing or tail feathers of a bird
    --3 is a kind of feather, plume, plumage
    --3 is a part of wing
    --3 has particulars: primary, primary feather, primary quill; tail feather
  4. quill, calamus, shaft - the hollow shaft of a feather
    --4 is a kind of rib
    --4 is a part of feather, plume, plumage
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